Difference Between Prokaryotic And Eukaryotic Cell Pdf Viewer
Prokaryotes are organisms made up of cells that lack a cell nucleus or any membrane-encased organelles. This means the genetic material DNA in prokaryotes is not bound within a nucleus. Additionally, the DNA is less structured in prokaryotes than in eukaryotes. Aladdin hardlock usb emulator software. In prokaryotes, DNA is a single loop.
Abstract
Eukaryotes have long been thought to have arisen by evolving a nucleus, endomembrane, and cytoskeleton. In contrast, it was recently proposed that the first complex cells, which were actually proto-eukaryotes, arose simultaneously with the acquisition of mitochondria. This so-called symbiotic association hypothesis states that eukaryotes emerged when some ancient anaerobic archaebacteria (hosts) engulfed respiring alpha-proteobacteria (symbionts), which evolved into the first energy-producing organelles. Therefore, the intracellular compartmentalization of the energy-converting metabolism that was bound originally to the plasma membrane appears to be the key innovation towards eukaryotic genome and cellular organization. The novel energy metabolism made it possible for the nucleotide synthetic apparatus of cells to be no longer limited by subsaturation with substrates and catalytic components. As a consequence, a considerable increase has occurred in the size and complexity of eukaryotic genomes, providing the genetic basis for most of the further evolutionary changes in cellular complexity. On the other hand, the active uptake of exogenous DNA, which is general in bacteria, was no longer essential in the genome organization of eukaryotes. The mitochondrion-driven scenario for the first eukaryotes explains the chimera-like composition of eukaryotic genomes as well as the metabolic and cellular organization of eukaryotes.
Full Text
Selected References
These references are in PubMed. This may not be the complete list of references from this article.
- Adam RD. The biology of Giardia spp. Microbiol Rev. 1991 Dec;55(4):706–732.[PMC free article] [PubMed] [Google Scholar]
- Akhmanova A, Voncken F, van Alen T, van Hoek A, Boxma B, Vogels G, Veenhuis M, Hackstein JH. A hydrogenosome with a genome. Nature. 1998 Dec 10;396(6711):527–528. [PubMed] [Google Scholar]
- Andersson SG, Kurland CG. Reductive evolution of resident genomes. Trends Microbiol. 1998 Jul;6(7):263–268. [PubMed] [Google Scholar]
- Battista JR. Against all odds: the survival strategies of Deinococcus radiodurans. Annu Rev Microbiol. 1997;51:203–224. [PubMed] [Google Scholar]
- Biderre C, Pagès M, Méténier G, Canning EU, Vivarès CP. Evidence for the smallest nuclear genome (2.9 Mb) in the microsporidium Encephalitozoon cuniculi. Mol Biochem Parasitol. 1995 Nov;74(2):229–231. [PubMed] [Google Scholar]
- Brown JR, Doolittle WF. Archaea and the prokaryote-to-eukaryote transition. Microbiol Mol Biol Rev. 1997 Dec;61(4):456–502.[PMC free article] [PubMed] [Google Scholar]
- Bui ET, Bradley PJ, Johnson PJ. A common evolutionary origin for mitochondria and hydrogenosomes. Proc Natl Acad Sci U S A. 1996 Sep 3;93(18):9651–9656.[PMC free article] [PubMed] [Google Scholar]
- Cavalier-Smith T. Eukaryotes with no mitochondria. Nature. 326(6111):332–333. [PubMed] [Google Scholar]
- Cavalier-Smith T, Chao EE. Molecular phylogeny of the free-living archezoan Trepomonas agilis and the nature of the first eukaryote. J Mol Evol. 1996 Dec;43(6):551–562. [PubMed] [Google Scholar]
- Clark CG, Roger AJ. Direct evidence for secondary loss of mitochondria in Entamoeba histolytica. Proc Natl Acad Sci U S A. 1995 Jul 3;92(14):6518–6521.[PMC free article] [PubMed] [Google Scholar]
- Doolittle RF. Microbial genomes opened up. Nature. 1998 Mar 26;392(6674):339–342. [PubMed] [Google Scholar]
- Doolittle WF. A paradigm gets shifty. Nature. 1998 Mar 5;392(6671):15–16. [PubMed] [Google Scholar]
- Doolittle WF. You are what you eat: a gene transfer ratchet could account for bacterial genes in eukaryotic nuclear genomes. Trends Genet. 1998 Aug;14(8):307–311. [PubMed] [Google Scholar]
- Embley TM, Hirt RP. Early branching eukaryotes? Curr Opin Genet Dev. 1998 Dec;8(6):624–629. [PubMed] [Google Scholar]
- Embley TM, Martin W. A hydrogen-producing mitochondrion. Nature. 1998 Dec 10;396(6711):517–519. [PubMed] [Google Scholar]
- Errington J. Bacillus subtilis sporulation: regulation of gene expression and control of morphogenesis. Microbiol Rev. 1993 Mar;57(1):1–33.[PMC free article] [PubMed] [Google Scholar]
- Ferat JL, Michel F. Group II self-splicing introns in bacteria. Nature. 1993 Jul 22;364(6435):358–361. [PubMed] [Google Scholar]
- Fonstein M, Haselkorn R. Physical mapping of bacterial genomes. J Bacteriol. 1995 Jun;177(12):3361–3369.[PMC free article] [PubMed] [Google Scholar]
- Fuerst JA, Webb RI. Membrane-bounded nucleoid in the eubacterium Gemmata obscuriglobus. Proc Natl Acad Sci U S A. 1991 Sep 15;88(18):8184–8188.[PMC free article] [PubMed] [Google Scholar]
- Germot A, Philippe H, Le Guyader H. Evidence for loss of mitochondria in Microsporidia from a mitochondrial-type HSP70 in Nosema locustae. Mol Biochem Parasitol. 1997 Aug;87(2):159–168. [PubMed] [Google Scholar]
- Gray MW, Burger G, Lang BF. Mitochondrial evolution. Science. 1999 Mar 5;283(5407):1476–1481. [PubMed] [Google Scholar]
- Gupta RS. Protein phylogenies and signature sequences: A reappraisal of evolutionary relationships among archaebacteria, eubacteria, and eukaryotes. Microbiol Mol Biol Rev. 1998 Dec;62(4):1435–1491.[PMC free article] [PubMed] [Google Scholar]
- Gupta RS, Aitken K, Falah M, Singh B. Cloning of Giardia lamblia heat shock protein HSP70 homologs: implications regarding origin of eukaryotic cells and of endoplasmic reticulum. Proc Natl Acad Sci U S A. 1994 Apr 12;91(8):2895–2899.[PMC free article] [PubMed] [Google Scholar]
- Hashimoto T, Sánchez LB, Shirakura T, Müller M, Hasegawa M. Secondary absence of mitochondria in Giardia lamblia and Trichomonas vaginalis revealed by valyl-tRNA synthetase phylogeny. Proc Natl Acad Sci U S A. 1998 Jun 9;95(12):6860–6865.[PMC free article] [PubMed] [Google Scholar]
- Holland HD. Evidence for life on Earth more than 3850 million years ago. Science. 1997 Jan 3;275(5296):38–39. [PubMed] [Google Scholar]
- Horner DS, Hirt RP, Kilvington S, Lloyd D, Embley TM. Molecular data suggest an early acquisition of the mitochondrion endosymbiont. Proc Biol Sci. 1996 Aug 22;263(1373):1053–1059. [PubMed] [Google Scholar]
- Jensen KF, Pedersen S. Metabolic growth rate control in Escherichia coli may be a consequence of subsaturation of the macromolecular biosynthetic apparatus with substrates and catalytic components. Microbiol Rev. 1990 Jun;54(2):89–100.[PMC free article] [PubMed] [Google Scholar]
- Kasting JF. Earth's early atmosphere. Science. 1993 Feb 12;259(5097):920–926. [PubMed] [Google Scholar]
- Keeling PJ, Doolittle WF. Evidence that eukaryotic triosephosphate isomerase is of alpha-proteobacterial origin. Proc Natl Acad Sci U S A. 1997 Feb 18;94(4):1270–1275.[PMC free article] [PubMed] [Google Scholar]
- Kelman Z, O'Donnell M. DNA replication: enzymology and mechanisms. Curr Opin Genet Dev. 1994 Apr;4(2):185–195. [PubMed] [Google Scholar]
- Kondrashov AS, Crow JF. Haploidy or diploidy: which is better? Nature. 1991 May 23;351(6324):314–315. [PubMed] [Google Scholar]
- Lawrence JG, Ochman H. Molecular archaeology of the Escherichia coli genome. Proc Natl Acad Sci U S A. 1998 Aug 4;95(16):9413–9417.[PMC free article] [PubMed] [Google Scholar]
- Margolin W. A green light for the bacterial cytoskeleton. Trends Microbiol. 1998 Jun;6(6):233–238. [PubMed] [Google Scholar]
- Margulis L. Archaeal-eubacterial mergers in the origin of Eukarya: phylogenetic classification of life. Proc Natl Acad Sci U S A. 1996 Feb 6;93(3):1071–1076.[PMC free article] [PubMed] [Google Scholar]
- Martin W, Müller M. The hydrogen hypothesis for the first eukaryote. Nature. 1998 Mar 5;392(6671):37–41. [PubMed] [Google Scholar]
- Moreira D, Lopez-Garcia P. Symbiosis between methanogenic archaea and delta-proteobacteria as the origin of eukaryotes: the syntrophic hypothesis . J Mol Evol. 1998 Nov;47(5):517–530. [PubMed] [Google Scholar]
- Moyer CL, Morita RY. Effect of growth rate and starvation-survival on cellular DNA, RNA, and protein of a psychrophilic marine bacterium. Appl Environ Microbiol. 1989 Oct;55(10):2710–2716.[PMC free article] [PubMed] [Google Scholar]
- Reiner DS, Shinnick TM, Ardeshir F, Gillin FD. Encystation of Giardia lamblia leads to expression of antigens recognized by antibodies against conserved heat shock proteins. Infect Immun. 1992 Dec;60(12):5312–5315.[PMC free article] [PubMed] [Google Scholar]
- Rivera MC, Jain R, Moore JE, Lake JA. Genomic evidence for two functionally distinct gene classes. Proc Natl Acad Sci U S A. 1998 May 26;95(11):6239–6244.[PMC free article] [PubMed] [Google Scholar]
- Roger AJ, Svärd SG, Tovar J, Clark CG, Smith MW, Gillin FD, Sogin ML. A mitochondrial-like chaperonin 60 gene in Giardia lamblia: evidence that diplomonads once harbored an endosymbiont related to the progenitor of mitochondria. Proc Natl Acad Sci U S A. 1998 Jan 6;95(1):229–234.[PMC free article] [PubMed] [Google Scholar]
- Schopf JW. Microfossils of the Early Archean Apex chert: new evidence of the antiquity of life. Science. 1993 Apr 30;260:640–646. [PubMed] [Google Scholar]
- Schulz HN, Brinkhoff T, Ferdelman TG, Mariné MH, Teske A, Jorgensen BB. Dense populations of a giant sulfur bacterium in Namibian shelf sediments. Science. 1999 Apr 16;284(5413):493–495. [PubMed] [Google Scholar]
- Seto S, Miyata M. Cell reproduction and morphological changes in Mycoplasma capricolum. J Bacteriol. 1998 Jan;180(2):256–264.[PMC free article] [PubMed] [Google Scholar]
- Sogin ML. Early evolution and the origin of eukaryotes. Curr Opin Genet Dev. 1991 Dec;1(4):457–463. [PubMed] [Google Scholar]
- Vellai T, Takács K, Vida G. A new aspect to the origin and evolution of eukaryotes. J Mol Evol. 1998 May;46(5):499–507. [PubMed] [Google Scholar]
- Wheeler RT, Shapiro L. Bacterial chromosome segregation: is there a mitotic apparatus? Cell. 1997 Mar 7;88(5):577–579. [PubMed] [Google Scholar]
- Woese CR. Bacterial evolution. Microbiol Rev. 1987 Jun;51(2):221–271.[PMC free article] [PubMed] [Google Scholar]
- Woese C. The universal ancestor. Proc Natl Acad Sci U S A. 1998 Jun 9;95(12):6854–6859.[PMC free article] [PubMed] [Google Scholar]
All living organisms can be sorted into one of two groups depending on the fundamental structure of their cells: the prokaryotes and the eukaryotes. Prokaryotes are organisms made up of cells that lack a cell nucleus or any membrane-encased organelles. Eukaryotes are organisms made up of cells that possess a membrane-bound nucleus that holds genetic material as well as membrane-bound organelles.
Understanding Cells and Cell Membranes
The cell is a fundamental component of our modern definition of life and living things. Cells are regarded as the basic building blocks of life and are used in the elusive definition of what it means to be 'alive.'
Cells keep chemical processes tidy and compartmentalized so individual cell processes do not interfere with others and the cell can go about its business of metabolizing, reproducing, etc. To achieve this, cell components are enclosed in a membrane which serves as a barrier between the outside world and the cell's internal chemistry. The cell membrane is a selective barrier, meaning that it lets some chemicals in and others out. In so doing it maintains the chemical balance necessary for the cell to live.
The cell membrane regulates the crossing of chemicals in and out of the cell in three ways including:

Differences Between Prokaryotic And Eukaryotic Organisms
- Diffusion (the tendency of solute molecules to minimize concentration and thus move from an area of higher concentration towards an area of lower concentration until concentrations equalize)
- Osmosis (the movement of solvent across a selective boundary in order to equalize the concentration of a solute that is unable to move across the boundary)
- Selective transport (via membrane channels and membrane pumps)
Prokaryotes
Prokaryotes are organisms made up of cells that lack a cell nucleus or any membrane-encased organelles. This means the genetic material DNA in prokaryotes is not bound within a nucleus. In addition, the DNA is less structured in prokaryotes than in eukaryotes: in prokaryotes, DNA is a single loop while in Eukaryotes DNA is organized into chromosomes. Most prokaryotes are made up of just a single cell (unicellular) but there are a few that are made of collections of cells (multicellular).
Scientists have divided the prokaryotes into two groups, the Bacteria, and the Archaea. Some bacteria, including E Coli, Salmonella, and Listeria, are found in foods and can cause disease; others are actually helpful to human digestion and other functions. Archaea were discovered to be a unique life form which is capable of living indefinitely in extreme environments such as hydrothermal vents or arctic ice.
A typical prokaryotic cell might contain the following parts:
- Cell wall: the membrane surrounding and protecting the cell
- Cytoplasm: all of the material inside a cell except the nucleus
- Flagella and pili: protein-based filaments found on the outside of some prokaryotic cells
- Nucleoid: a nucleus-like region of the cell where genetic material is kept
- Plasmid: a small molecule of DNA that can reproduce independently
Eukaryotes
Eukaryotes are organisms made up of cells that possess a membrane-bound nucleus (that holds DNA in the form of chromosomes) as well as membrane-bound organelles. Eukaryotic organisms may be multicellular or single-celled organisms. All animals are eukaryotes. Other eukaryotes include plants, fungi, and protists.
A typical eukaryotic cell is surrounded by a plasma membrane and contains many different structures and organelles with a variety of functions. Examples include the chromosomes (a structure of nucleic acids and protein which carry genetic information in the form of genes), and the mitochondria (often described as the 'powerhouse of the cell').